t is the probable range of dates for the panin / hominin divergence (i.e., when is it
likely that the earliest member of the hominin tribe appeared?)
the panin-hominin divergence occurred between 5 and 8 million years ago
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Sahelanthropus tchadensis?
Chad, 7 m.y.a. (estimate based on biostratigraphy,
surprisingly old) cranial capacity estimated 320 to
380 cm3 (chimpanzee size) mixture of panin-like
and hominin-like features: huge browridges, sagittal
crest, large muscle attachments, yet smallish vertical
face with front teeth that are very un-apelike?no
postcranial remains, so bipedal status unknown
(hominin status inferred from face and teeth)
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Orrorin tugenensis?
Ethiopia, 6 m.y.a.
mostly dental fragments, cranial capacity unknown;
some lower leg bones that apparently indicate
bipedalism
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Ardipithecus ramidus?
Ethiopia (Aramis), 4.4 m.y.a.; fragmentary cranial
remains, but cranial capacity probably between 300
and 350 cc; thin molar enamel (unusual for early
hominins); bipedal pelvis, forward position of
foramen magnum, forelimb not weight bearing?all
indicating bipedalism, but divergent big toe
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Australopithecus anamensis?
northern Kenya (Kanapoi), 4.2 ? 3.9 m.y.a.
cranial capacity unknown
primitive features including large canines; the
most primitive Australopithecine discovered so
far, but still clearly a small-brained biped with
thick molar enamel
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Australopithecus afarensis?
Ethiopia (Hadar), Chad, Laetoli (Tanzania)
ranging from 3.9 ? 3.0 m.y.a.
cranial capacity averaging 420 cm3 based on
extensive specimens (more than 60
individuals)
very primitive: large canines, parallel teeth
rows, but definitely bipedal
high degree of sexual dimorphism (males 5?,
females 3.5?)
?Lucy,? Laetoli footprints; a strong candidate
for the ancestor of all later hominins, based on
currently available evidence
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Australopithecus africanus?
southern Africa, 2.5 ? 2.0 m.y.a. (no
Australopithecine fossils have been found in
East Africa after 3.0 m.y.a.)
cranial capacity averaging 440 cm3; smallbodied,
large-toothed bipeds
the ?Taung child?
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Paranthropus robustus and Paranthropus boisei?
South Africa, 2.0 ? 1.3 m.y.a.
cranial capacity averaging 520 cm3
massive jaws and teeth, large broad faces, large
sagittal crest
specialized diet (seeds, nuts, and roots)
East Africa (Olduvai Gorge, Tanzania), 2.3 ?
1.0 m.y.a.
cranial capacity averaging 520 cm3
massive jaws and teeth, large broad faces, large
sagittal crest
specialized diet (seeds, nuts, and roots)
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Homo habilis?
Olduvai Gorge, Tanzania, East Africa, 1.8 m.y.a.
average cranial capacity 630 cm3
smaller and narrower teeth, slightly larger body size
compared to the Australopithecines
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Homo erectus?
Homo erectus lived from 1,800,000 to 200,000 (or even
later) years ago, during the Pleistocene (?Ice Age?)
East Africa 1.8 m.y.a.
Georgia (Dmanisi) 1.75 m.y.a. (possibly an earlier form
of Homo)
Java 1.6 m.y.a. (possibly surviving to 25,000 y.a.)
China as early as 670,000 y.a.
European presence of Homo (in Spain) as early as 1.2
m.y.a. years ago was probably not Homo erectus, but
another species (perhaps Homo antecessor)
age cranial capacity 900 cm3 (with a range from 700
cm3 to 1,250 cm3)
large brow ridges (supraorbital tori), heavy cranial bones
postorbital constriction
receding forehead and chin, first prominent hominin nose
long, low skull vault, with maximum breadth low on the
skull
pentagonal skull shape from rear, shovel-shaped incisors
larger body than early Homo (average 5?6?, > 100 lbs),
with considerable sexual dimorphism (males average 6?
tall)
Nariokotome Boy (a.k.a. Turkana Boy, Lake Turkana,
Kenya, 1984), 1.6 m.y.a., 8-12 years old
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Homo heidelbergensis?
Africa, Asia, & Europe
850,000 years ago ? 200,000 years ago
average cranial capacity 1,200 cm3
increased parietal breadth
skull no longer pentagonal
occipital bun
pronounced brow ridges (compared to Homo sapiens)
decreased cranial and postcranial robusticity (compared
to Homo erectus)
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Homo sapiens neanderthalensis?
Europe and southwest Asia only; 130,000 ? 30,000
years ago
average cranial capacity 1,520 cm3
(largest brain of any hominin that?s ever lived; possible
adaptation to extreme cold)
large, long, low cranium, bulging at the sides
prominent brow ridges, projecting face, large nasal
aperture
very robust postcranially?barrel-chested, heavily
muscled
What are the dates and biological characteristics (i.e., diagnostic cranial and postcranial
traits) of Homo sapiens sapiens?
Africa 195,000 y.a.
Southwest Asia 130,000 y.a., East Asia 40,000 y.a.,
Southeast Asia 45,000
Australia ~50,000 y.a., Europe 35,000 y.a.
average cranial capacity 1,400 cm3
high, rounded cranial vault; vertical forehead; distinct
chin; small brow ridges; less robust post-cranially
little or no prognathism (the extent to which the lower
face protrudes in front of the upper face in profile)
smaller teeth evolved because of specialized tools; chin
evolved to buttress lower jaw (good example of
biocultural evolution)
What is known for certain about the exact human phylogeny from the Pliocene and
Pleistocene?
Homo heidelbergensis definitely descended from Homo erectus (although some
regional populations of Homo erectus may have become extinct)
Modern humans definitely descended from Homo heidelbergensis (although some
regional populations of Homo heidelbergensis may have become extinct)